[86] Kushata JNT, Périquet S, Tarakini T, Muzamba M, Mafuwa B, Loveridge AJ, Macdonald DW, Fritz H & Valeix M (2018) Drivers of diurnal rest site selection by spotted hyaenas. Journal of Zoology 304: 132-140. pdf [86]

[85] Valls-Fox H, De Garine-Wichatitsky M, Fritz H & Chamaillé-Jammes S (2018) Resource depletion versus landscape complementation: habitat selection by a multiple central place forager. Landscape Ecology 33: 127-140. pdf [85]


[84] Blanchard P, Pays O & Fritz H (2017) Ticks or lions: trading between allogrooming and vigilance maternal care. Animal Behaviour 129: 269-279. pdf [84]

[83] Guerbois C & Fritz H (2017) Patterns and perceived sustainability of provisioning ecosystem services on the edge of a protected area in times of crisis. Ecosystem Services 28: 196-206. pdf [83]

[82] Loveridge AJ, Valeix M, Elliot NB & Macdonald DW (2017) The landscape of anthropogenic mortality: how African lions respond to spatial variation in risk. Journal of Applied Ecology 54: 815-825. pdf [82]

[81] Miguel E, Grosbois V, Fritz H, Caron A, de Garine-Wichatitsky M, Nicod F, Loveridge AJ, Stapelkamp B, Macdonald DW & Valeix M (2017) Drivers of foot-and-mouth disease in cattle at wild/domestic interface: insights from farmers, buffalo and lions. Diversity and Distributions 23: 1018-1030. pdf [81]

[80] Msiteli-Shumba S, Kativu S & Hulot F (2017) Influence of environmental variables on plankton community composition in permanent and temporal pans in and around Hwange National Park, Zimbabwe. Transactions of the Royal Society of South Africa 72: 266-279. pdf[80]

[79] Schweitzer C, Gaillard T, Guerbois C, Fritz H & Petit O (2017) Participant profiling and pattern of crop-foraging in chacma baboon (Papio hamadryas Ursinus) in Zimbabwe: why does investigating age-sex classes matters? International Journal of Primatology 38: 207-223. pdf [79]

[78] Tshipa A, Valls-Fox H, Fritz H, Collins K, Sebele L, Mundy P & Chamaillé-Jammes S (2017) Partial migration links local surface-water management to large-scale elephant conservation in the world’s largest transfrontier conservation area. Biological Conservation 215: 46-50. pdf [78]


[77] Barnier F, Duncan P, Fritz H, Blanchard P, Rubenstein DI & Pays O (2016) Between-gender differences in vigilance do not necessarily lead to differences in foraging-vigilance tradeoffs. Oecologia 181: 757-768. pdf [77]

[76] Chamaillé-Jammes S, Charbonnel A, Dray S, Madzikanda H & Fritz H (2016) Spatial distribution of a large herbivore community at waterholes: an assessment of its stability over years in Hwange National Park, Zimbabwe. PLoS ONE 11(4): e0153639.   pdf [76]

[75] Courbin N, Loveridge AJ, Macdonald DW, Fritz H, Valeix M, Makuwe ET & Chamaillé-Jammes S (2016) Reactive responses of zebras to lion encounters shape their predator-prey space game at large scale. Oikos 125: 829-838. pdf [75]

[74] Ferry N, Dray S, Fritz H & Valeix M (2016) Interspecific interference competition at the resource patch scale: do large herbivores spatially avoid elephants while accessing water? Journal of Animal Ecology 85: 1574-1585. pdf [74]

[73] Loveridge AJ, Valeix M, Chapron G, Davidson Z, Mtare G & Macdonald DW (2016) Conservation of large predator populations: demographic and spatial responses of African lions to the intensity of trophy hunting. Biological Conservation 204: 247-254. pdf [73]

[72] Périquet S, Mapendere C, Revilla E, Banda J, Macdonald DW, Loveridge AJ & Fritz H (2016) A potential role for interference competition with lions in den selection and attendance by spotted hyaenas. Mammalian Biology 81: 227-234. pdf [72]

[71] Sirot E, Renaud PC, Pays O (2016) How competition and predation shape patterns of waterhole use by herbivores in arid ecosystems. Animal Behaviour 118: 19-26. pdf [71]

[70] Wigley BJ, Slingsby JA, Díaz S, Bond WJ, Fritz H & Coetsee C (2016) Leaf traits of African woody savanna species across climate and soil fertility gradients: evidence for conservative versus acquisitive resource-use strategies. Journal of Ecology 104: 1357-1369. pdf [70]


[69] Crosmary W, Chamaillé-Jammes S, Mtare G, Côté SD & Fritz H (2015) Decline of sable antelope in one of its key conservation areas: the greater Hwange ecosystem, Zimbabwe. African Journal of Ecology 53: 94-205. pdf [69]

[68] Crosmary WG, Côté SD & Fritz H (2015) The assessment of the role of trophy hunting in wildlife conservation. Animal Conservation 18: 136-137. pdf [68]

[67] Crosmary WG, Côté S & Fritz H (2015) Does trophy hunting matter to long-term trends in African herbivores of different dietary guilds? Animal Conservation 18: 117-130. pdf [67]

[66] Grange S, Barnier F, Duncan P, Gaillard J-M, Valeix M, Ncube H, Périquet S & Fritz H (2015) Demography of plains zebras (Equus quagga) under heavy predation. Population Ecology 57: 201-214. pdf [66]

[65] Périquet S, Valeix M, Claypole J, Drouet-Hoguet N, Salnicki J, Revilla E & Fritz H (2015) Spotted hyaenas switch their foraging strategy as a response to changes in intraguild interactions with lions. Journal of Zoology 297: 245-254. pdf [65]

[64] Périquet S, Fritz H & Revilla E (2015) Large carnivore interactions and coexistence: the Lion King and the Hyaena Queen. Biological Reviews 90: 1197-1214. pdf [64]

[63] van der Meer E, Fritz H & Pays O (2015) Anti-predator behaviour of kudu and impala in response to mimicked African wild dog presence: do age and sex matter? Behaviour 152: 1209-1228. pdf [63]

[62] van der Meer E, Rasmussen GSA & Fritz H (2015) Using an energetic cost-benefit approach to identify ecological traps: the case of the African wild dog. Animal Conservation 18: 359-366. pdf [62]


[61] Barnier F, Valeix M, Duncan P, Chamaillé-Jammes S, Barre P, Loveridge AJ, Macdonald DW & Fritz H (2014) Diet quality in a wild grazer declines under the threat of an ambush predator. Proceedings of the Royal Society B 281: 20140446. pdf [61]

[60] Benhamou S, Valeix M, Chamaillé-Jammes S, Macdonald DW & Loveridge AJ (2014) Movement-based analysis of interactions in African lions. Animal Behaviour 90: 171-180. pdf [60]

[59] Elliot NB, Valeix M, Macdonald DW & Loveridge AJ (2014) Social relationships affect dispersal timing revealing a delayed infanticide in African lions. Oikos 123: 1049-1056. pdf [59]

[58] Miguel E, Boulinier T, de Garine-Wichatitsky M, Caron A, Fritz H & Grosbois V (2014) Characterizing African tick communities at a wild-domestic interface using repeated sampling protocols and models. Acta Tropica 138: 5-14. pdf [58]

[57] Pays O, Ekori A & Fritz H (2014) On the advantages of mixed-species groups: impalas adjust their vigilance when associated with larger prey herbivores. Ethology 120: 1207-1216. pdf [57]

[56] Tarakini T, Crosmary WG, Fritz H & Mundy P (2014) Flight behavioural responses to sport hunting by two African herbivores. South Africa Journal of Wildlife Research 44: 76-83. pdf [56]

[55] van der Meer E, Fritz H, Blinston P & Rasmussen GSA (2014) Ecological trap in the buffer zone of the protected area: effects of indirect anthropogenic mortality on the African wild dog Lycaon pictus. Oryx 48: 285-293. pdf [55]

[54] van der Meer E, Rasmussen GSA, Muvengwi J & Fritz H (2014) Foraging costs, hunting success and its implications for African wild dogs (Lycaon pictus) conservation inside and outside a protected area. African Journal of Ecology 52: 69-76. pdf [54]


[53] Chamaillé-Jammes S, Mtare G, Makuwe E & Fritz H (2013) African elephants adjust speed in response to surface-water constraint on foraging during the dry season. PloS One 8: e59164. pdf [53]

[52] Crosmary WG, Loveridge AJ, Ndaimani H, Le Bel S, Booth V, Côté SD & Fritz H (2013) Trophy hunting in Africa: long-term trends in antelope horn size. Animal Conservation 16: 648-660. pdf [52]

[51] Davidson Z, Valeix M, Van Kesteren F, Loveridge AJ, Hunt JE, Murindagomo F & Macdonald DW (2013) Seasonal diet and prey preference of the African lion in a waterhole-driven semi-arid savanna. PLoS ONE 8(2): e55182. pdf [51]

[50] de Garine Wichatitsky M, Miguel E, Mukamuri E, de Garine I, Ancelcius J, Pfukenyi D & Caron A (2013) Coexisting with wildlife in Transfrontier Conservation Areas in Zimbabwe: cattle owners’ awareness of disease risks and perception of the role played by wildlife. Comparative Immunology, Microbiology and Infectious Disease 36: 321-332. pdf [50]

[49] Favreau FR, Pays O, Goldizen AW & Fritz H (2013) Short-term behavioural responses of impalas in simulated antipredator and social contexts. PLoS ONE 8: e84970. pdf [49]

[48] Guerbois C, Dufour AB, Mtare G & Fritz H (2013) Insights for integrated conservation from attitudes of peole toward protected areas near Hwange National Park, Zimbabwe. Conservation Biology 27: 844-855. pdf [48]

[47] Miguel E, Grosbois V, Caron A, Boulinier T, Fritz H, Cornélis D, Foggin C, Makaya PV, Tshabalala PT & de Garine-Wichatitsky M (2013) Contacts and foot and mouth disease transmission from wild to domestic bovines in Africa. Ecosphere 4: 1-32. pdf [47]

[46] Riotte-Lambert L, Benhamou S & Chamaillé-Jammes S (2013) Periodicity analysis of movement recursions. Journal of Theoretical Biology 317: 238-243. pdf [46]

[45] Seeber PA, Duncan P, Fritz H & Gandswindt A (2013) Androgen changes and flexible rutting behaviour in male giraffes. Biology Letters 9(5): e20130396. pdf [45]

[44] van der Meer E, Mpofu J, Rasmussen GSA, Muvengwi J & Fritz H (2013) Characteristics of African wild dog natal dens selected under different interspecific predation pressures. Mammalian Biology 78: 336-343. pdf [44]


[43] Barnier F, Grange S, Gandswindt A, Ncube H & Duncan P (2012) Inter-birth interval in zebras is longer following the birth of male foals than after female foals. Acta Oecologica 42: 11-15. pdf [43]

[42] Crosmary WG, Valeix M, Fritz H, Madzikanda H & Côté SD (2012) African ungulates and their drinking problems: hunting and predation risks constrain access to water. Animal Behaviour 83: 145-153. pdf [42]

[41] Crosmary WG, Makumbe P, Côté S & Fritz H (2012) Vulnerability to predation and water constraints limit behavioural adjustments of ungulates in response to hunting risk. Animal Behaviour 83: 1367-1376. pdf [41]

[40] Davidson Z, Valeix M, Loveridge AJ, Hunt JE, Johnson PJ, Madzikanda H & Macdonald DW (2012) Environmental determinants of habitat and kill site selection in a large carnivore: scale matters. Journal of Mammalogy 93: 667-685. pdf [40]

[39] Guerbois C, Chapanda E & Fritz H (2012) Combining multi-scale socio-ecological approaches to understand the susceptibility of subsistence farmers to elephant crop raiding on the edge of a protected area. Journal of Applied Ecology 49: 1149-1158. pdf [39]

[38] Pays O, Blanchard P, Valeix M, Chamaillé-Jammes S, Duncan P, Périquet S, Lombard M, Ncube G, Tarakini T, Makuwe E & Fritz H (2012) Detecting predators and locating competitors while foraging: an experimental study of a medium-sized herbivore in an African savanna. Oecologia 169: 419-430. pdf [38]

[37] Pays O, Sirot E & Fritz H (2012) Collective vigilance in the greater kudu: towards a better understanding of synchronisation patterns. Ethology 118: 1-9. pdf [37]

[36] Périquet S, Todd-Jones L, Valeix M, Stapelkamp B, Elliot N, Wijers M, Pays O, Fortin D, Madzikanda H, Fritz H, Macdonald DW & Loveridge AJ (2012) Influence of immediate predation risk by lions on the vigilance of prey of different body size. Behavioral Ecology 23: 970-976. pdf [36]

[35] Seeber PA, Ndlovu HT, Duncan P & Ganswindt A (2012) Grazing behaviour of the giraffe in Hwange National Park, Zimbabwe. African Journal of Ecology 50: 247-250. pdf [35]

[34] Valeix M, Loveridge AJ & Macdonald DW (2012) Influence of prey dispersion on territory and group size of African lions: a test of the resource dispersion hypothesis. Ecology 93: 2490-2496. pdf [34]

[33] van der Meer E, Pays O & Fritz H (2012) The effect of simulated African wild dog presence on anti-predator behaviour of kudu and impala. Ethology 118: 1018-1027. pdf [33]


[32] Davidson Z, Valeix M, Loveridge AJ, Madzikanda H & Macdonald DW (2011) Socio-spatial behaviour of an African lion population following perturbation by sport hunting. Biological Conservation 144: 114-121. pdf [32]

[31] Fritz H, Loreau M, Chamaillé-Jammes S, Valeix M & Clobert J (2011) A food web perspective on large herbivore community limitation. Ecography 34: 196-202. pdf [31]

[30] Ncube H, Duncan P, Grange S, Cameron EZ, Barnier F & Gandswindt A (2011) Patterns of faecal 20-oxopregnane and oestrogen concentrations during pregnancy in wild plains zebra mares. General Comparative Endocrinology 172: 358-362. pdf [30]

[29] Valeix M, Chamaillé-Jammes S, Loveridge AJ, Davidson Z, Hunt JE, Madzikanda H & Macdonald DW (2011) Understanding patch departure rules for large carnivores: lion movements support a patch-disturbance hypothesis. The American Naturalist 178: 269-275. pdf [29]

[28] Valeix M, Fritz H, Sabatier R, Murindagomo F, Cumming D & Duncan P (2011) Elephant-induced structural changes in the vegetation and habitat selection by large herbivores in an African savanna. Biological Conservation 144: 902-912. pdf [28]

[27] Valeix M (2011) Temporal dynamics of dry-season water-hole use by large African herbivores in two years of contrasting rainfall in Hwange National Park, Zimbabwe. Journal of Tropical Ecology 27: 163-170. pdf [27]

[26] van der Meer E, Moyo M, Rasmussen GSA & Fritz H (2011) An empirical and experimental test of the risk and costs of kleptoparasitism for African wild dogs (Lycaon pictus) inside and outside a protected area. Behavioral Ecology 22: 985-992. pdf [26]


[25] Martin J, Chamaillé-Jammes S, Nichols JD, Fritz H, Hines JE, Fonnesbeck CJ, MacKenzie DI & Bailey LL (2010) Simultaneous modeling of habitat suitability, occupancy, and relative abundance: African elephants in Zimbabwe. Ecological Applications 20: 1173-1182. pdf [25]

[24] Midgley J, Lawes M & Chamaillé-Jammes S (2010) Savanna woody plant dynamics, the role of fire and herbivory, separately and synergistically. Australian Journal of Botany 58: 1-11. pdf [24]

[23] Périquet S, Valeix M, Loveridge AJ, Madzikanda H, Macdonald DW & Fritz H (2010) Individual vigilance of African herbivores while drinking: the role of immediate predation risk and context. Animal Behaviour 79: 665-671. pdf [23]

[22] Valeix M, Loveridge AJ, Davidson Z, Madzikanda H, Fritz H & Macdonald DW (2010) How key habitat features influence large terrestrial carnivore movements: waterholes and African lions in semi-arid savanna of north-western Zimbabwe. Landscape Ecology 25: 337-351. pdf [22]


[21] Chamaillé-Jammes S, Valeix M, Bourgarel M, Murindagomo F & Fritz H (2009) Seasonal density estimates of common large herbivores in Hwange National Park, Zimbabwe. African Journal of Ecology 47: 804-808. pdf [21]

[20] Chamaillé-Jammes S, Fritz H & Madzikanda H (2009) Piosphere contribution to landscape heterogeneity: a case-study of remote-sensed woody cover in a high elephant density landscape. Ecography 32: 871-880. pdf [20]

[19] Chamaillé-Jammes S & Fritz H (2009) Precipitation-NDVI relationships in eastern and southern African savannas vary along a precipitation gradient. International Journal of Remote Sensing 30: 3409-3422. pdf [19]

[18] Loveridge AJ, Valeix M, Davidson Z, Murindagomo F, Fritz H & Macdonald DW (2009) Changes in home range size of African lions in relation to pride size and prey biomass in a semi-arid savanna. Ecography 32: 953-962. pdf [18]

[17] Valeix M, Loveridge AJ, Chamaillé-Jammes S, Davidson Z, Murindagomo F, Fritz H & Macdonald DW (2009) Behavioral adjustments of African herbivores to predation risk by lions: spatiotemporal variations influence habitat use. Ecology 90: 23-30. pdf [17]

[16] Valeix M, Fritz H, Loveridge AJ, Davidson Z, Hunt JE, Murindagomo F & Macdonald DW (2009) Does the risk of encountering lions influence African herbivore behaviour at waterholes? Behavioral Ecology and Sociobiology 63: 1483-1494. pdf [16]

[15] Valeix M, Fritz H, Canévet V, Le Bel S & Madzikanda H (2009) Do elephants prevent other African herbivores from using waterholes in the dry season? Biodiversity and Conservation 18: 569-576. pdf [15]


[14] Blanchard P, Sabatier R & Fritz H (2008) Within-group spatial position and vigilance: the case of impalas (Aepyceros melampus) with a controlled food supply. Behavioral Ecology and Sociobiology 12: 1863-1868. pdf [14]

[13] Blanchard P & Fritz H (2008) Seasonal variation in rumination parameters of free-ranging impalas (Aepyceros melampus). Wildlife Biology 14: 372-378. pdf [13]

[12] Chamaillé-Jammes S, Fritz H, Valeix M, Murindagomo F & Clobert J (2008) Resource variability, aggregation and direct density dependence in an open context: the local regulation of an African elephant population. Journal of Animal Ecology 77: 135-144. pdf [12]

[11] Valeix M, Fritz H, Chamaillé-Jammes S, Bourgarel M & Murindagomo F (2008) Fluctuations in abundance of large herbivore populations: insights into the influence of dry season rainfall and elephant numbers from long-term data. Animal Conservation 11: 391-400. pdf [11]

[10] Valeix M, Fritz H, Matsika R, Matsvimbo F & Madzikanda H (2008) The role of water abundance, thermoregulation, perceived predation risk, and interference competition in water access by African herbivores. African Journal of Ecology 46: 402-410. pdf [10]


[9] Blanchard P & Fritz H (2007) Induced or routine vigilance while foraging. Oikos 116: 1603-1608. pdf [9]

[8] Chamaillé-Jammes S, Valeix M & Fritz H (2007) Managing heterogeneity in elephant distribution: interactions between elephant population density and surface-water availability. Journal of Applied Ecology 44: 625-633. pdf [8]

[7] Chamaillé-Jammes S, Valeix M & Fritz H (2007) Elephant management: why can’t we throw out the babies with the artificial bathwater? Diversity and Distributions 13: 663-665. pdf [7]

[6] Chamaillé-Jammes S, Fritz H & Holdo R (2007) Spatial relationship between elephant and sodium concentration of water disappears as density increases in Hwange National Park, Zimbabwe. Journal of Tropical Ecology 23: 725-728. pdf [6]

[5] Chamaillé-Jammes S, Fritz H & Murindagomo F (2007) Climate-driven fluctuations in surface-water availability and the buffering role of artificial pumping in an African savanna: potential implication for herbivore dynamics. Austral Ecology 32: 740-748. pdf [5]

[4] Chamaillé-Jammes S, Fritz H & Murindagomo F (2007) Detecting climate changes of concern in highly variable environments: quantile regressions reveal that droughts worsen in Hwange National Park, Zimbabwe. Journal of Arid Environments 71: 321-326. pdf [4]

[3] Valeix M, Chamaillé-Jammes S & Fritz H (2007) Interference competition and temporal niche shifts: elephants and herbivore communities at waterholes. Oecologia 153: 739-748. pdf [3]

[2] Valeix M, Fritz H, Dubois S, Kanengoni K, Alleaume S & Saïd S (2007) Vegetation structure and ungulate abundance over a period of increasing elephant abundance in Hwange National Park, Zimbabwe. Journal of Tropical Ecology 23: 87-93. pdf [2]


[1] Chamaillé-Jammes S, Fritz H & Murindagomo F (2006) Spatial patterns of the NDVI-rainfall relationship at the seasonal and interannual time-scales in an African savanna. International Journal of Remote Sensing 27: 5185-5200. pdf [1]